i've been using the following groups of Trichocereus
BRI
CUZ
MAC
PAC
SCO
VAL
TER
Those familiar with the epithets will have no problem figuring out the groups
it is my hypothesis that the original ancestors of the groups are those hailing from the region of Bolivia and Argentina and that over time the forms we know came to be realized and grown
I believe that the VAL and TER groups are the older groups and that the San Pedro complex arose from the introduction of alleles into VAL and TER populations via hybridization with echinopsis like forms and subsequent selection over time
I believe that the alleles for short spines were introduced into the groups this way, as were other alleles including those for the prostrate growth of the MAC group.
I also believe that this is the origin of the alleles in the CUZ group
In terms of age I believe the TER and VAL group to be the oldest, with the SCO and BRI groups being next and with the MAC, CUZ and PAC groups arising over time
I believe that the initial spread of the groups was facilitated by the transportation and propagation of seeds and that selection and asexual propagation occurred after-wards, I also believe that this occurred over the course of more than 10,000 years and that sub-populations have been inbreeding and stabilizing over this time to create a reticulated phylogeny.
In terms of the 7 major sub-groups I believe there to be several intermediate populations, particularly for the MAC,PAC and CUZ groups
I believe the MAC and PAC groups to originate as cultigens, the CUZ to represent a stable feral group descending from a cultigen, a cultigen which still has extant examples in some regions
I believe that the allele diversity found in these groups was primarily the result of human selection and emphasis upon seed grown plants that were the result of random crosses and that the flow of alleles was not linear in any geographic sense due to the extreme periods of time and human migrations involved in the area.
I also believe that the cosmology and mythology of the precolumbian peoples holds major clues relative to the flow of people from the Tiwanako area to northern coastal Peru over 10,000 years ago. The story of Manco Cápac and his emergence at Isla Del Sol relates to this.
i also believe that in horticulture we have many forms with a unique combination of alleles which are not found in the andean region, that there at present exists more diversity in the group as cultivated in modern times than there did in over 10,000 years of cultivation and use by the andean peoples. It is my belief that this, in concert with the reticulated phylogeny, obfuscates the direction of flow in terms of the origin of the cultigens in modern use, but that the groups of VAL and TER will be demonstrated to be ancestral to the MAC, PAC, BRI and CUZ group.
I would like to see detailed sampling of extant populations in terms of alleles and diversity, and mitochondrial studies of the major groups and their sub-populations.
this is my theory at present and it is relative and subject to change